Through lectures, readings, class exercises and discussion, this class provides a critical survey of Lifecourse epidemiology and its application to the public's health. In this class we will systematically survey both seminal and current literature on lifecourse epidemiology along with key related topics ranging from cellular to societal function. We consider the debates and controversies that ensued following publication of many of the groundbreaking papers and the subsequent revisions and evolution of the field in the context of these controversies. The second half of the class surveys the current state of science of Lifecourse epidemiology. We conclude this course by focusing on the growing literature on paternal influences within the Lifecourse perspective as well as opportunities for promoting health over the generations.